Bacillus subtilis: a plant-growth promoting rhizobacterium that also impacts biotic stress

Bacillus subtilis: a plant-growth promoting rhizobacterium that also impacts biotic stress

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SAUDI JOURNAL OF BIOLOGICAL SCIENCES Volume 26, Issue 6, September 2019, Pages 1291-1297 https://doi.org/10.1016/j.sjbs.2019.05.004Get rights and content ABSTRACT Plants encounter many biotic agents, such as viruses, bacteria, nematodes, weeds, and arachnids. These entities induce biotic stress in their hosts by disrupting normal metabolism, and as a result, limit plant growth and/or are the cause of plant mortality. Some biotic agents, however, interact symbiotically or synergistically with their host plants. Some microbes can be beneficial to plants and perform the same role as chemical fertilizers and pesticides, acting as a biofertilizer and/or biopesticide. Plant growth promoting rhizobacteria (PGPR) can significantly enhance plant growth and represent a mutually helpful plant-microbe interaction. _Bacillus_ species are a major type of rhizobacteria that can form spores that can survive in the soil for long period of time under harsh environmental conditions. Plant growth is enhanced by PGPR through the induction of systemic resistance, antibiosis, and competitive omission. Thus, the application of microbes can be used to induce systemic resistance in plants against biotic agents and enhance environmental stress tolerance. _Bacillus subtilis_ exhibits both a direct and indirect biocontrol mechanism to suppress disease caused by pathogens. The direct mechanism includes the synthesis of many secondary metabolites, hormones, cell-wall-degrading enzymes, and antioxidants that assist the plant in its defense against pathogen attack. The indirect mechanism includes the stimulation of plant growth and the induction of acquired systemic resistance. _Bacillus subtilis_ can also solubilize soil P, enhance nitrogen fixation, and produce siderophores that promote its growth and suppresses the growth of pathogens. _Bacillus subtilis_ enhances stress tolerance in their plant hosts by inducing the expression of stress-response genes, phytohormones, and stress-related metabolites. The present review discusses the activity of _B. subtilis_ in the rhizosphere, its role as a root colonizer, its biocontrol potential, the associated mechanisms of biocontrol and the ability of _B. subtilis_ to increase crop productivity under conditions of biotic and abiotic stress. KEYWORDS Rhizobacteria _Bacillus subtilis_ Biocontrol potential Biocontrol mechanism Biotic stress Abiotic stress ABBREVIATIONS PGPR plant growth promoting rhizobacteria ACC 1-aminocyclopropane-1-carboxylate deaminase VOCs volatile organic compounds PGP plant growth promotion ISR induced systemic resistance LPs lipopeptides JA jasmonic acid PAL phenylalanine ammonialyase POD peroxidase PPO polyphenol oxidase SOD superoxide dismutase GA3 gibberellic acid IAA indole acetic acid ABA abscisic acid CITED BY (0) * Peer review under responsibility of King Saud University. © 2019 The Authors. Production and hosting by Elsevier B.V. on behalf of King Saud University.

SAUDI JOURNAL OF BIOLOGICAL SCIENCES Volume 26, Issue 6, September 2019, Pages 1291-1297 https://doi.org/10.1016/j.sjbs.2019.05.004Get rights and content ABSTRACT Plants encounter many


biotic agents, such as viruses, bacteria, nematodes, weeds, and arachnids. These entities induce biotic stress in their hosts by disrupting normal metabolism, and as a result, limit plant


growth and/or are the cause of plant mortality. Some biotic agents, however, interact symbiotically or synergistically with their host plants. Some microbes can be beneficial to plants and


perform the same role as chemical fertilizers and pesticides, acting as a biofertilizer and/or biopesticide. Plant growth promoting rhizobacteria (PGPR) can significantly enhance plant


growth and represent a mutually helpful plant-microbe interaction. _Bacillus_ species are a major type of rhizobacteria that can form spores that can survive in the soil for long period of


time under harsh environmental conditions. Plant growth is enhanced by PGPR through the induction of systemic resistance, antibiosis, and competitive omission. Thus, the application of


microbes can be used to induce systemic resistance in plants against biotic agents and enhance environmental stress tolerance. _Bacillus subtilis_ exhibits both a direct and indirect


biocontrol mechanism to suppress disease caused by pathogens. The direct mechanism includes the synthesis of many secondary metabolites, hormones, cell-wall-degrading enzymes, and


antioxidants that assist the plant in its defense against pathogen attack. The indirect mechanism includes the stimulation of plant growth and the induction of acquired systemic resistance.


_Bacillus subtilis_ can also solubilize soil P, enhance nitrogen fixation, and produce siderophores that promote its growth and suppresses the growth of pathogens. _Bacillus subtilis_


enhances stress tolerance in their plant hosts by inducing the expression of stress-response genes, phytohormones, and stress-related metabolites. The present review discusses the activity


of _B. subtilis_ in the rhizosphere, its role as a root colonizer, its biocontrol potential, the associated mechanisms of biocontrol and the ability of _B. subtilis_ to increase crop


productivity under conditions of biotic and abiotic stress. KEYWORDS Rhizobacteria _Bacillus subtilis_ Biocontrol potential Biocontrol mechanism Biotic stress Abiotic stress ABBREVIATIONS


PGPR plant growth promoting rhizobacteria ACC 1-aminocyclopropane-1-carboxylate deaminase VOCs volatile organic compounds PGP plant growth promotion ISR induced systemic resistance LPs


lipopeptides JA jasmonic acid PAL phenylalanine ammonialyase POD peroxidase PPO polyphenol oxidase SOD superoxide dismutase GA3 gibberellic acid IAA indole acetic acid ABA abscisic acid


CITED BY (0) * Peer review under responsibility of King Saud University. © 2019 The Authors. Production and hosting by Elsevier B.V. on behalf of King Saud University.